Quarterly report pursuant to Section 13 or 15(d)

Balance Sheets Details - Property and Equipment, net (Details)

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Balance Sheets Details - Property and Equipment, net (Details) - USD ($)
$ in Thousands
Sep. 30, 2019
Dec. 31, 2018
Property, Plant and Equipment [Line Items]    
Property and equipment $ 39,920 $ 37,852
Less: accumulated depreciation and amortization (33,679) (33,093)
Property and equipment, net 6,241 4,759
Laboratory equipment [Member]    
Property, Plant and Equipment [Line Items]    
Property and equipment 23,266 21,328
Leasehold improvements [Member]    
Property, Plant and Equipment [Line Items]    
Property and equipment 10,899 10,359
Computer equipment and software [Member]    
Property, Plant and Equipment [Line Items]    
Property and equipment 3,806 3,954
Office equipment and furniture [Member]    
Property, Plant and Equipment [Line Items]    
Property and equipment 1,461 1,272
Construction in progress [Member]    
Property, Plant and Equipment [Line Items]    
Property and equipment $ 488 $ 939