Annual report pursuant to Section 13 and 15(d)

Balance Sheets Details - Property and Equipment, net (Details)

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Balance Sheets Details - Property and Equipment, net (Details) - USD ($)
$ in Thousands
12 Months Ended
Dec. 31, 2019
Dec. 31, 2018
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment $ 39,533 $ 37,852
Less: accumulated depreciation and amortization (33,251) (33,093)
Property and equipment, net 6,282 4,759
Laboratory equipment    
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment 23,561 21,328
Equipment retired during period 1,000 300
Leasehold improvements    
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment 10,804 10,359
Computer equipment and software    
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment 3,016 3,954
Office equipment and furniture    
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment 1,461 1,272
Construction in progress    
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment $ 691 $ 939