Annual report pursuant to Section 13 and 15(d)

Balance Sheet Details - Property and Equipment, net (Details)

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Balance Sheet Details - Property and Equipment, net (Details) - USD ($)
$ in Thousands
12 Months Ended
Dec. 31, 2015
Dec. 31, 2014
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment $ 35,324 $ 37,288
Less: accumulated depreciation and amortization (32,215) (31,452)
Impairment of laboratory equipment 0 (1,841)
Property and equipment, net 3,109 3,995
Laboratory equipment    
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment 20,503 23,002
Leasehold improvements    
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment 10,369 9,773
Computer equipment and software    
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment 3,271 3,262
Office equipment and furniture    
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment 1,178 1,227
Construction in progress    
Property, Plant and Equipment, Net, by Type [Abstract]    
Property and equipment $ 3 $ 24